A NEW EPIGEAN FALSE SCORPION : RONCUS SUMADIJAE N . SP . ( NEOBISIIDAE , PSEUDOSCORPIONES ) FROM THE BALKAN PENINSULA ( WESTERN SERBIA )

— A new endemic epigean species from the village of Adžina Livada, nr. Kragujevac, Mts. Gledićke Planine, western Serbia, is erected, described and thoroughly illustrated. Its main morphometric characters and important diagnostic features are analyzed and compared to the two closest congeners, Roncus ivanjicae B. Ćurčić, and R. golijae B. Ćurčić from western Serbia, respectively.


INTRODUCTION
A major breakthrough in the study of both epigean and cavernicolous species of the genus Roncus L. Koch, 1873 in Serbia has been carried out during the past years by Ćurčić (1992a, b), Ćurčić and Dimitrijević (2009), Ćurčić et al. (1993, 2004, 2006, 2010c), and Dimitrijević (2000).The known number of taxa assigned to this genus in Serbia has risen from seven in 1993 to the present 32.Of these, 21 species are cave dwelling and 11 inhabit leaf-litter and humus.The majority of these species are relict and endemic forms for Serbia, i.e. for the Balkan Peninsula.
Female genital area: unknown.
Cheliceral galea in the form of a small and low hyaline tubercle (Fig. 8).Cheliceral palm bears 6 long setae, movable finger with one seta.Both cheliceral fingers with small close-set and apically rounded denticles, which diminish proximally and are eventually replaced with rounded teeth.Movable chelal finger with 2 or 3 distal larger teeth and a series of 12 teeth which diminish in size proximally; fixed finger with a row of 13 or 14 close-set and triangular, but apically rounded denticles which diminish proximally.Galeal seta inserted, basal to larger teeth on the movable finger (Fig. 8).The flagellum has eight blades, one small blade proximally and seven blades twice this length, more or less in pairs, distally.The most distal member of the series are curved but all, to some extent, are pinnate on two sides (Fig. 5).
Apex of pedipalpal coxa bears five long and acuminate setae.Trochanter and tibia are short, other pedipalpal articles are smooth and elongate (Fig. 3).Pedipalpal femur with interior and lateral granulations, tibia smooth, and pedipalpal chela and pedipalpal chelal palm with some interior and lateral tubercles (Fig. 3).There are no small setae proximal to eb and esb; instead, 5 or 8 such setae exist dorsal to the two trichobothria (Fig. 1).Chelal fingers of almost equal size.Fixed chelal finger with 47-50 close-set teeth; movable finger teeth with 42-44 such teeth.Chelal fingers longer than chelal palm; fixed chelal finger of nearly equal length as pedipalpal femur, pedipalpal femur shorter or longer than carapace (Table 1).
Trichobothriotaxy: ib and isb on the palm of the chela; the fixed finger carries a further 6 trichobothria (et, est, esb, eb, it, and ist); the movable finger bears 4 trichobothria (t, st, sb, and b) (Fig. 1).
Pedal coxae II and III bear some coxal spines.Tibia IV, basitarsus IV, and telotarsus IV each with a single tactile seta.
Measurements and morphometric ratios are presented in Table 1.
Differential diagnosis -Roncus sumadijae n. sp. is easily distinguished from R. ivanjicae B. Ćurčić, and R. golijae B. Ćurčić in the form of appendages, setation of tergites and sternites, body size, linear measurements and ratio of different characters, as well as in the type locality (Table 1; Ćurčić, 1988).
Distribution -Since the distribution center of the genus Roncus is found in southern Europe, it is probable that it evolved there.This assumption is further supported by the discovery of several Roncus-related genera in the caves of southwestern Europe and by the discovery of new living genera from the Dinaric karst and Maritime Alps, both of which have  (Ćurčić, in preparation).The discovery of these "proton roncids" justifies the assumption regarding the great age and autochthonous origin of the Roncus stock in the Dinaric region, as well as in the Mediterranean area as whole (Ćurčić, 1972, 1984, 1988, 1992a, b;Ćurčić and Beron, 1981;Ćurčić et al., 1993, 2004, 2010a, b, c, d, e, f, g;2011a, b, c, d, e, f, g, h;Hadži, 1937).It seems probable that the ancestral population was broadly distributed over the ancient continent of Laurasia.Subsequently, with a break up of this supercontinent, North American and Eurasian genera and species evolved differently.As already mentioned, on the Balkan Peninsula some ancient taxa have survived; however, the majority of Roncus species probably originated during the Tertiary period.Finally, the Roncus-related taxa then, are of different age, integrating the species of Laurasian, Paleo-Mediterranean and Proto-Balkan origin (Ćurčić, 1986; Ćurčić et al., 2004).
more primitive characteristics compared to Roncus