ON SOME NEW CAVE-DWELLING GROUND BEETLES (COLEOPTERA: CARABIDAE: TRECHINI) FROM EASTERN SERBIA

The following new cavernicolous ground beetle taxa are described from three caves in eastern Serbia: Duvalius (Paraduvalius) trifunovici sp. n., from the Mandina Pećina Cave, village of Zlot, near Bor, Kučajske Planine Mts., D. (P.) rtanjensis sp. n., from the Golema Porica Pit, Mt. Rtanj, and Glabroduvalius gen. n., G. tupiznicensis sp. n., from the Gornja Lenovačka Pećina Cave, village of Lenovac, near Zaječar, Mt. Tupižnica. The new taxa are easily distinguished from related organisms. All important morphological features have been listed, along with the diagnoses and illustrations of the taxa. The new taxa are relicts and endemics of eastern Serbia and probably belong to old phyletic lineages of Tertiary or even pre-Tertiary origin.

In recent years, several new trechine genera have been described from Serbia -Serboduvalius Ćurčić, Pavićević & Ćurčić, 2001, Rascioduvalius Ćurčić, Brajković, Mitić & Ćurčić, 2003, Curcicia Ćurčić & Brajković, 2003, and Javorella Ćurčić, Brajković & Ćurčić, 2003(Ćurčić et al., 2001, 2003b, 2003c;Ćurčić and Brajković, 2003).Certain regions of eastern, western, and southern Serbia remain unexplored, where some new trechine taxa can be expected to be found in the future.Some field trips in eastern Serbia, organized by the Institute of Zoology, University of Belgrade -Faculty of Biology in 2012, resulted in the discovery of four new trechine taxa: a genus and three species.Descriptions and diagnoses of the new trechine taxa are given in the present study.The diagnoses of Du-valius (Paraduvalius) trifunovici sp.n. and D. (P.) rtanjensis sp n. are based on a thorough analysis of the type series of three males each collected in the Mandina Pećina Cave, Kučajske Planine Mts., near Bor and the Golema Porica Pit, Mt.Rtanj, respectively, while the diagnoses of Glabroduvalius gen.n. and G. tupiznicensis sp.n. are based on a thorough analysis of the type series of two males and a female collected in the Gornja Lenovačka Pećina Cave, Mt.Tupižnica, near Zaječar.

MATERIALS AND METHODS
The ground beetle specimens were analyzed in the laboratories of the Institute of Zoology, University of Belgrade -Faculty of Biology, Belgrade, Serbia.The genitalia were removed from the bodies and fixed on microscope slides in a medium composed of Canada balsam and xylol.Afterwards the beetles were glued onto paper labels and then analyzed as dry specimens.All taxonomically important morphological characters were studied for comparison.A Carl Zeiss Stemi 2000 binocular stereomicroscope and Carl Zeiss Axioskop 40 microscope with a Canon Pow-erShot A80 digital camera attached were used in this study.Additionally, Canon PowerShot SX 130 IS and Canon EOS 400D digital cameras were used for photographing whole specimens.) stankovitchi by its smaller size (4.47-4.94mm vs. 5.00-6.20 mm), length/width ratio of the head (wider than long vs. equal length and width) and the pronotum (wider than long vs. longer than wide), shape of the posterior pronotal angles (somewhat obtuse angled vs. almost right-angled), length/width ratio of the elytra (1.69 times as long as wide vs. 2.00 times as long as wide), number of well-depressed elytral striae (three inner ones vs. four inner ones), position of the first pair of elytral discal setae (on third elytral striae vs. on fourth elytral interstriae), and size and shape of the aedeagus (medium-sized, almost forming a right angle somewhat after its basal third, then almost straight, median lobe slightly narrowing towards the apex, less abruptly narrowing apically, basal bulb rounded vs. massive, elongated, abruptly bent around its basal third, convex dorsally in that part, then straight, median lobe of a constant width towards the apex, more abruptly narrowing apically, basal bulb with a basal narrowing), and copulatory piece (as long as twothirds of aedeagus length, widening medially, then narrowing towards the apex, apex acute vs. as long as three-fourths of aedeagus length, of a constant width, apically narrowing, apex obtuse) (Jeannel, 1923(Jeannel, , 1928;;present study).

RESULTS AND DISCUSSION
Duvalius (Paraduvalius) trifunovici sp.n. is easily distinguished from D. (P.) winkleri by its bigger size (4.47-4.94mm vs. 4.20 mm), length/width ratio of the head (wider than long vs. equal length and width), form of the eyes (ellipsoid, with a darkened border vs. in the form of a whitish elongated ring, with no trace of pigment), length of the antennae (exceeding slightly over the mid-elytra level vs. almost reaching the mid-elytra level), length/width ratio of the pronotum (wider than long vs. longer than wide), shape of the posterior pronotal angles (somewhat obtuse angled vs. somewhat acute), presence of hairs on the pronotum (present vs. absent), position of the first pair of pronotal setae (at the anterior third of pronotal length vs. slightly below the anterior fourth of pronotal length), shape of the shoulders (slightly elevated vs. fairly prominent), position of the first pair of elytral discal setae (around the anterior fourth of elytral length, at the level of the fourth humeral setae vs. around the anterior fifth of elytral length, above the level of the third humeral setae), size of the legs as compared to the body size (long vs. short), and size and shape of the aedeagus (medium-sized, almost forming a right angle somewhat after the basal third, then almost straight, median lobe slightly narrowing towards the apex, abruptly narrowing apically, basal bulb massive, rounded vs. massive, elongated, curved over the whole of its length, with a sub-apical widening, then gradually narrowing apically, apex obtuse, basal bulb medium-sized), and copulatory piece (as long as two-thirds of aedeagus length, elongated and thin, apically pointed, with no spines vs. as long as a third of aedeagus length, short and wide, triangular, with a large number of fine spines apically) (Jeannel, 1923(Jeannel, , 1928;;present study).
Duvalius (Paraduvalius) trifunovici sp.n. is easily distinguished from D. (P.) rtanjensis sp.n. by its bigger size (4.47-4.94 mm vs. 4.29-4.41 mm), body color (reddish-brown vs. light brown), size and form of the eyes (0.12 mm long, 0.06 mm wide, ellipsoid vs. 0.10 mm long, 0.03 mm wide, lenticular), number of the ommatidia (15-20 vs. 6-8), length of the antennae (exceeding slightly over the middle of the elytra vs. almost reaching the middle of the elytra), shape of the posterior pronotal angles (somewhat obtuse-angled vs. almost right-angled), shape of the posterior pronotal margin (straight vs. slightly concave medially), position of the first pair of pronotal setae (at the anterior third of pronotal length vs. around the anterior fourth of pronotal length), shape of the shoulders (slightly elevated, rounded vs. lowered, obtuse-angled), distance between the humeral setae and the position (the longest distance between the second and third setae, the shortest distance between the third and fourth setae; the second seta right next to the marginal furrow, the first seta slightly away from the furrow vs. humeral setae equidistant; first and second setae located at the edge of the marginal furrow), length/width ratio of the elytra (1.69 times longer than wide vs. 1.73 times longer than wide), the position of the first pair of elytral discal setae (around the anterior fourth of elytral length, at the level of the fourth humeral setae vs. around the anterior fifth of elytral length, at the level of the second humeral setae), and size and shape of the aedeagus (longer, almost forming a right angle somewhat after the basal third, then almost straight, median lobe slightly narrowing towards the apex, basal bulb massive vs. shorter, regularly curved, slightly convex dorsally in the middle, basal bulb medium-sized), copulatory piece (as long as two-thirds of aedeagus length, unifid, apically pointed vs. slightly shorter than half of aedeagus length, bifid, apex in the form of two slightly pronounced lobes), and urite (slightly longer than the aedeagus, more elongated vs. scarcely shorter than the aedeagus, less elongated) (present study).
Head stout, rounded, slightly wider than long.Frontal furrows deep, complete, and arcuate.Hairs lacking, except for cheeks.Cheeks convex.Eyes reduced, small, 0.12 mm long and 0.06 mm wide, ellipsoid, with a darkened border, composed of 15-20 depigmented ommatidia.An arcuated preocular furrow present.Mentum tooth bifid.Antennae of moderate length, exceeding slightly over the middle of elytra.Antennomere II shorter than antennomere IV.Antennal article XI about three times as long as wide (Fig. 1).
Elytra elongated, ovoid, 1.69 times as long as wide, widest slightly below the mid level.Shoulders rounded, slightly elevated.Elytral apex rounded.Marginal furrows narrow and deep.Elytral disc slightly convex.First three inner striae developed, well depressed.Outer striae present in a form of more or less pronounced rows of points.Inner interstrial spaces convex.
Legs long and thin.Fore tibias with a longitudinal fissure each.Male protarsi I and II dilated (Fig. 1).
Chaetotaxy: Head with two pairs of supraorbital setae.Pronotum with two pairs of pronotal setae.The first pair located at the anterior third, while the second pair situated in posterior pronotal angles.Each humerus with four setae.The longest distance is between the second and third setae, the shortest distance is between the third and fourth.The second seta right next to the marginal furrow.First, third, and fourth setae away from the furrow, with the first one being closest to it, while the fourth one is farthest from it.Two discal setae on each elytron.The first pair of elytral discal setae situated on third striae, around the anterior fourth of elytral length, at the level of the fourth humeral setae.The second pair of elytral discal setae situated on third striae, somewhat below the mid-elytra level (Fig. 1).
Female genitalia unknown since no females were found.
Bionomy and distribution -The new species was found on the floor, under rocks and a piece of rotten wood, as well as in pitfall traps baited with rotten meat in the middle part of the Mandina Pećina Cave, village of Zlot, near Bor, Kučajske Planine Mts., eastern Serbia.For the time being, the species is known only from the type locality.
Remarks -The new species belongs to the "stankovitchi" group of species based on the shape of the elytra, presence of unpronounced shoulders (rounded, not angled), absence of laid hairs on the vertex, and presence of a longitudinal fissure on the fore tibias and the first elytral discal setae located below the level of the third humeral setae (Guéorguiev, 1971).5.00-6.20 mm), length/width ratio of the head (wider than long vs. as long as wide), shape of the pronotum (wider than long vs. longer than wide), position of the first pair of pronotal setae (around the anterior fourth of pronotal length vs. slightly above the anterior third of pronotal length), length/width ratio of the elytra (1.73 times as long as wide vs. 2.00 times as long as wide), shape of the shoulders (lowered vs. elevated), position of the first pair of elytral discal setae (around the anterior fifth of elytral length, on third striae, at the level of the second humeral setae vs. around the anterior fourth of elytral length, in fourth interstriae, below the level of the third humeral setae), number of well-depressed striae (three inner ones vs. four inner ones), and size and shape of the aedeagus (medium-sized, curved, slightly convex dorsally in the middle part, apex acute, basal bulb medium-sized, rounded vs. massive, elongated, abruptly bent around the basal third, convex dorsally in that part, then straight, median lobe of a constant width towards the apex, abruptly narrowing apically, basal bulb massive, with a basal narrowing) and copulatory piece (slightly shorter than half of aedeagus length, bifid, apex in the form of two lobes vs. as long as three-fourths of aedeagus length, unifid, with a rounded apex) (Jeannel, 1923(Jeannel, , 1928;;present study).

DUVALIUS (PARADUVALIUS) RTANJENSIS
Duvalius (Paraduvalius) rtanjensis sp.n. is easily distinguished from D. (P.) winkleri by its bigger size (4.29-4.41mm vs. 4.20 mm), length/width ratio of the head (wider than long vs. equal length and width), form of the eyes (lenticular, with a darkened border vs. in the form of a whitish, elongated ring, with no trace of pigment), length/width ratio of the prono-tum (wider than long vs. longer than wide), shape of the posterior pronotal angles (almost right-angled vs. somewhat acute) and pronotal margin (slightly concave medially vs. straight), presence of hairs on the pronotum (present vs. absent), length/width ratio of the elytra (1.73 times as long as wide vs. 1.67 times as long as wide), shape of the shoulders (obtuse-angled, lowered vs. rounded, fairly prominent, elevated), position of the humeral setae (first and second setae located at the edge of the marginal furrow, third and fourth setae slightly away from the furrow, with the fourth one slightly farther away from it than the third vs. all humeral setae located next to the marginal furrow) and second pair of elytral discal setae (slightly below the mid-elytra level vs. slightly above the mid-elytra level), size of the legs as compared to the body size (long vs. short), and size and shape of the aedeagus (medium-sized, curved, slightly convex dorsally in the middle part, narrowing apically, apex acute, basal bulb larger vs. massive, elongated, curved over the whole of its length, with a sub-apical widening, then narrowing apically, apex obtuse, basal bulb smaller) and copulatory piece (slightly shorter than half of aedeagus length, elongated, with no spines, bifid, apex in the form of two lobes vs. as long as a third of aedeagus length, triangular, short, wide, with a large number of fine spines apically, unifid) (Jeannel, 1923(Jeannel, , 1928;;present study).
Head stout, rounded, slightly wider than long.Frontal furrows deep, complete, and arcuate.Cheeks rounded, covered with tiny hairs.Eyes reduced, small, 0.10 mm long and 0.03 mm wide, lenticular, with a darkened border, the anterior margin convex, the posterior one flattened, composed of 6-8 depig-mented ommatidia.An arcuated pre-ocular furrow present.Mentum tooth bifid.Antennae of moderate length, almost reaching the middle of the elytra.Antennomere II shorter than antennomere IV.Antennal article XI about three times as long as wide (Fig. 5).
Pronotum small, wider than long, slightly wider than head, sub-cordate, widest at the anterior third, narrowing towards the base.Anterior pronotal margin slightly concave.Anterior pronotal angles prominent and rounded.Lateral pronotal margins rounded anteriorly and somewhat concave posteriorly.Posterior pronotal angles prominent, sharp, almost right-angled.Posterior pronotal margin slightly concave medially.Pronotal disc convex, with a median furrow.Marginal furrows narrow and shallow.Basal foveae wide, deep, and slightly punctate (Fig. 5).
Elytra elongated, ovoid, 1.73 times as long as wide, widest slightly below the mid-level.Shoulders lowered, obtuse-angled.Elytral apex rounded.Marginal furrows narrow and deep.Elytral disc slightly convex.First three inner striae developed, well depressed.Fourth striae incomplete, while other striae present in the form of rows of points.Inner interstrial spaces convex (Fig. 5).
Legs long and thin.Fore tibias with a longitudinal fissure each.Male protarsi I and II dilated (Fig. 5).
Chaetotaxy: Head with two pairs of supraorbital setae.Pronotum with two pairs of pronotal setae.The first pair located around the anterior fourth, while the second pair situated in posterior pronotal angles.Each humerus with four setae.Humeral setae equidistant.First and second setae located at the edge of the marginal furrow, while third and fourth setae slightly away from the furrow, with the fourth one slightly farther from it than the third.With two discal setae on each elytron.The first pair of elytral discal setae situated around the anterior fifth of elytral length, on third striae, at the level of the second humeral setae.The second pair of elytral discal setae situated on third striae, slightly below the mid-elytra level.
Female genitalia unknown since no females were found.
Bionomy and distribution -The new species was found under rocks on the floor and in pitfall traps at the end of one of the tunnels of the Golema Porica Pit, Mt.Rtanj, eastern Serbia.For the time being, the species is known only from the type locality.

Remarks -
The new species belongs to the "winkleri" group of species based on the shape of the elytra, presence of unpronounced shoulders (rounded, not angled), absence of laid hairs on the vertex, and presence of a longitudinal fissure on the fore tibias and the first elytral discal setae located above the level of the third humeral setae (Guéorguiev, 1971).
Diagnosis -Glabroduvalius gen.n. clearly differs from the following phenetically close genera from the Balkan Peninsula: Serboduvalius (from southwestern Serbia and eastern Montenegro), Rascioduvalius (from western Serbia), Curcicia (from eastern Serbia), Javorella (from southwestern and western Ser-bia), and Duvalius (a variable genus with a wider distribution that needs further revision; because of this, the comparison is made with the type species Duvalius raymondi Delarouzée, 1859) (Ćurčić et al., 2001, 2003b, 2003c;Ćurčić and Brajković, 2003).However, there are numerous distinctions between the new genus and the listed genera.Glabroduvalius gen.n. clearly differs from Serboduvalius in the body length (shorter vs. longer), form of the eyes (in the form of a darkened line vs. flattened and reduced), presence/ absence of the ommatidia (absent vs. present), body hairs (present vs. absent), and hairs on the cheeks (present vs. absent), shape of the shoulders (lowered, obtuse-angled, pointed vs. slightly lowered, obtuse, rounded), distance between the humeral setae and the position (the longest distance between the first and second setae, the shortest distance between the second and third; the first seta located on the inner edge of the marginal furrow, the second seta right next to the inner edge of the furrow, third and fourth setae located away from the furrow, with the fourth one slightly farther away from it than the third vs. the longest distance between the second and third setae, distances between setae I and II, as well as between setae III and IV are equal; the first seta located in the marginal furrow, the second on the inner edge of the furrow, the third and fourth setae slightly away from the furrow, both of these setae equidistant from the outer elytral edge), presence/absence of the elytral striae (absent vs. present), number of elytral discal setae (two vs. three in most cases), length/width ratio of the first male protarsomere (slightly longer than wide vs. twice as long as wide), size and shape of the aedeagus (small, regularly curved, abruptly narrowing apically vs. medium-sized, in most cases not regularly curved, sub-apically gradually narrowing towards the apex), number of setae on the paramerae (three vs. four), and shape of the copulatory piece (scarcely bifid vs. unifid in most cases) (Ćurčić et al., 2001; present study).
Glabroduvalius gen.n. clearly differs from Rascioduvalius in body length (shorter vs. longer), form of the cheeks (slightly convex vs. fairly convex) and eyes (in the form of a darkened line vs. small, elliptic, depigmented, and flattened), presence/absence of the ommatidia (absent vs. present), pre-ocular furrow (absent vs. present), and body hairs (present vs. absent), shape of the shoulders (lowered, obtuse angled, pointed vs. elevated, angulose, rounded), distance between the humeral setae and the position (the longest distance between the first and second setae, the shortest distance between the second and third; the first seta located on the inner edge of the marginal furrow, the second seta right next to the inner edge of the furrow, third and fourth setae located away from the furrow, with the fourth one slightly farther away from it than the third vs. distances between setae I and II, as well as between setae II and III, are equal, the shortest distance between the third and fourth setae; setae I and II, as well as III and IV equidistant from the outer elytral edge, but the distance is different in each of the two-setal pairs; first and second setae located on the inner edge of the marginal furrow, third and fourth setae slightly out of the furrow), presence/absence of the elytral striae (absent vs. present), shape of the legs (elongated vs. shortened) and aedeagus (less elongated, wider, median lobe slightly narrowing from the base towards the middle part, then slightly widening sub-apically and abruptly narrowing apically, basal bulb widening towards the base vs. more elongated, narrower, gradually narrowing towards the apex, basal bulb rounded), number of setae on the paramerae (three vs. four), and shape of the copulatory piece (scarcely bifid, wider vs. strongly bifid, narrower, tapering towards the apex, with two very long parallel lobes) (Jeannel, 1923(Jeannel, , 1928;;Ćurčić et al., 2003c, 2005; present study).
Glabroduvalius gen.n. clearly differs from Curcicia in body length (shorter vs. longer), presence/absence of the body hairs (present vs. absent), length/ width ratio of the head (more wide than long vs. equal length and width), presence/absence of hairs on the cheeks (present vs. absent), form of the cheeks (very slightly convex vs. more convex), presence/absence of the eyes (present, in the form of a darkened line vs. absent), length of the antennae (reaching between a third and a half of elytral length vs. exceeding over the mid-elytra level), shape of the shoulders (lowered, obtuse-angled, pointed vs. elevated, angulose, rounded), distance between the humeral setae and the position (the longest distance between the first and second setae, the shortest distance between second and third; the first seta located on the inner edge of the marginal furrow, the second seta right next to the furrow, third and fourth setae slightly away from the furrow, with the fourth one slightly farther away from it than the third vs. humeral setae equidistant, first and second setae located right next to the inner edge of the marginal furrow, third and fourth setae slightly away from the furrow), presence/absence of the elytral striae (absent vs. present) and longitudinal median fissure on the fore tibias (present vs. absent), shape of the aedeagus (less elongated, regularly curved, abruptly narrowing apically, apex straight, basal bulb medium-sized vs. more elongated, basally strongly curved, then almost straight, gradually narrowing from the sub-apical part towards the apex, apex curved upwards, basal bulb relatively small), number of setae on the paramerae (three vs. four), and shape of the copulatory piece (scarcely bifid, gutter-formed, wider vs. strongly bifid, not gutterformed, more elongated) (Pretner, 1963;Ćurčić and Brajković, 2003;present study).
Glabroduvalius gen.n. clearly differs from Javorella in body length (shorter vs. longer), length/ width ratio of the head (wider than long vs. longer than wide), form of the cheeks (very slightly convex vs. more convex), presence/absence of hairs on the cheeks (present vs. absent), form of the eyes (in the form of a darkened line vs. reduced, elliptic, pale, flattened, encircled with a slightly pigmented eye border), presence/absence of the pre-ocular furrow (absent vs. present) and ommatidia (absent vs. present), length of the antennae (shorter vs. longer), presence/ absence of the body hairs (present vs. absent), shape of the shoulders (lowered, obtuse-angled, pointed vs. elevated, distinctly marked, obtuse, rounded), distance between the humeral setae and the position (the shortest distance between the second and third setae; the first seta located on the inner edge of the marginal furrow, the second seta right next to the inner edge of the furrow, third and fourth setae located away from the furrow, the fourth one slightly farther away from it than the third vs. the shortest distance between the third and fourth setae; first and second setae located on the inner edge of the marginal furrow, third and fourth setae situated slightly out of the furrow, seta I closest to the elytral edge, seta II slightly farther from the edge, setae III and IV more spaced from the elytral edge and equidistant from it), presence/absence of the elytral striae (absent vs. present), number of setae on the paramerae (three vs. four), and shape of the copulatory piece (scarcely bifid vs. unifid) (Ćurčić et al., 2003a, 2003b; present study).
Glabroduvalius gen.n. is easily distinguished from Duvalius in body length (shorter vs. longer), presence/absence of the body hairs (present vs. absent), length/width ratio of the head (wider than long vs. longer than wide), form of the eyes (in the form of a darkened line vs. in the form of a whitish oval spot), presence/absence of the pre-ocular furrow (absent vs. present) and hairs on the cheeks (present vs. absent), length of the antennae (shorter vs. longer), length/width ratio of the pronotum (wider than long vs. longer than wide), shape of the posterior pronotal angles (acute-angled vs. almost right-angled) and shoulders (lowered, obtuse-angled, pointed vs. prominent, rounded), distance between the humeral setae and the position (the longest distance between the first and second setae, the shortest distance between the second and third; the first seta located on the edge of the marginal furrow, the second seta right next to the furrow, third and fourth setae slightly away from the furrow, with the fourth one slightly farther away from it than the third vs. all humeral setae right next to the marginal furrow, aggregated), presence/absence of the elytral striae (absent vs. present) and longitudinal median fissure on the fore tibias (present vs. absent), shape of the aedeagus (less elongated, with a pronounced sub-apical widening, abruptly narrowing apically, apex wider, basal bulb widening towards the base vs. more elongated, with a slight sub-apical widening, then gradually narrowing apically, apex narrower, basal bulb not widening towards the base), number of setae on the paramerae (three vs. four), and shape of the copulatory piece (scarcely bifid, wider vs. bifid, narrower) (Jeannel, 1928;present study).
Head stout, wider than long, slightly narrower than pronotum.Frontal furrows deep, complete, and arcuate.Cheeks very slightly convex, covered with tiny hairs.Eyes quite reduced, in the form of a darkened line, with no ommatidia.Pre-ocular furrow absent.Mentum tooth bifid.Antennae moderately elongate, reaching between a third and a half of elytral length (Fig. 9).Pronotum small, slightly wider than long, subcordate, covered with short rare hairs.Posterior angles prominent, acute-angled, sharp (Fig. 9).
Chaetotaxy: Head with two pairs of supraorbital setae.Pronotum with two pairs of pronotal setae.The first pair located around the anterior fourth, while the second pair situated in posterior pronotal angles.Each humerus with four setae.The longest distance between the first and second setae, the shortest distance between the second and third setae.The first seta located on the inner edge of the marginal furrow, the second seta right next to the furrow, third and fourth setae slightly away from the furrow, with the fourth one slightly farther away from it than the third.Two discal setae on each elytron.Both discal and umbilicate setae very long (Fig. 9).Aedeagus small, regularly curved.Basal bulb medium-sized.Paramerae with three setae each (Figs. 10 and 11).Copulatory piece medium-sized, about as long as half of aedeagus length, chitinized, gutter-shaped, scarcely bifid, the apex in the form of two slightly pronounced lobes (Fig. 11).Diagnosis -This is the only species in the genus so far, so comparison with other species could not be made for the time being.
Head stout, slightly wider than long.Frontal furrows deep, complete, and arcuate.Cheeks very slightly convex and covered with tiny hairs.Eyes quite reduced, in the form of a darkened line, with no ommatidia.Pre-ocular furrow absent.Antennae moderately elongate, reaching between a third and a half of elytral length.Antennomere II about of the same size as antennomere IV.Antennal article XI twice as long as wide (Fig. 9).Pronotum small, slightly wider than long, subcordate, widest around the anterior fourth, slightly wider than head, narrowing towards the base.Anterior pronotal margin slightly concave.Anterior pronotal angles prominent and rounded.Lateral pronotal margins rounded anteriorly and somewhat con-cave posteriorly.Posterior pronotal margin slightly concave in the middle.Posterior angles prominent, acute-angled, sharp.Pronotal disc convex, with a median furrow.Marginal furrows narrow and shal-low.Basal foveae wide, deep, and slightly punctate (Fig. 9).Elytra elongated, ovoid, 1.775 times as long as wide, widest slightly below the mid-level.Shoulders lowered, obtuse-angled, pointed.Elytral apex rounded.Elytral disc slightly convex.Marginal furrows narrow and deep.No developed striae, but instead of them rows of points occur (Fig. 9).
Chaetotaxy: Head with two pairs of supraorbital setae.Pronotum with two pairs of pronotal setae.The first pair located around the anterior fourth, the second pair situated in posterior pronotal angles.Each humerus with four setae.The longest distance between the first and second setae, the shortest distance between the second and third setae.The first seta located on the inner edge of the marginal furrow, the second seta right next to the furrow, third and fourth setae slightly away from the furrow, with the fourth one slightly farther away from it than the third.With two discal setae on each elytron.The first pair of elytral discal setae situated in fourth interstriae, about at the level of the anterior fourth of elytral length, slightly below the level of the third humeral setae.The second pair of elytral discal setae situated on third rows of points, somewhat below the midelytra level (Fig. 9).Both discal and umbilicate setae very long (Fig. 9).Aedeagus relatively small, regularly curved (lateral view) (Fig. 10).Median lobe gradually narrowing towards the sub-apical part, then somewhat widening, afterwards abruptly narrowing apically.Apex straight (lateral view) and rounded (dorsal view) (Figs. 10 and 11).Basal bulb medium-sized, widening towards the base (Fig. 10).Aedeagus scarcely bent to the left and gradually narrowing apically in dorsal view (Fig. 11).Paramerae moderately wide, narrowing towards the apices, with three setae each (Figs. 10 and 11).Copulatory piece about as long as half of aedeagus length, elongated, sub-parallel, chitinized, gutter-shaped, scarcely bifid, the apex in the form of two slightly pronounced lobes (Fig. 11).
Bionomy and distribution -The new species was found under rocks on the floor and in pitfall traps baited with rotten meat at the back of the Gornja Lenovačka Pećina Cave, village of Lenovac, near Zaječar, Mt.Tupižnica, eastern Serbia.For the time being, the species is known only from the type locality.

CONCLUSIONS
The new trechine taxa probably belong to old phyletic lineages of Tertiary or even pre-Tertiary origin (Guéorguiev, 1977).The taxa are both relicts and endemics of the Carpathian mountain chain in eastern Serbia.The endemic differentiation of the new taxa and relatives on the Balkan Peninsula was facilitated by the great Alpine Orogeny, paleoclimatic events, and subsequent evolution of the underground karstic relief, which yielded numerous new epigean and hypogean niches suitable for preservation of the old and autochthonous fauna (Ćurčić et al., 2012).Tupižnica, eastern Serbia.10 -holotype male, aedeagus (lateral view); 11 -holotype male, aedeagus (dorsal view); 12 -holotype male, abdominal sternite IX (urite); 13 -paratype female, genitalia.Scales = 0.20 mm.
of the Rock and Ice Society of Extreme Sports (Bor, Serbia).