NEW AND INTERESTING BRYOPHYTE RECORDS FOR THE FLORA OF SERBIA

During a revision of the bryophyte collections in the Herbarium of the Balkan Peninsula, Natural History Museum of Belgrade (BEO) and the Herbarium of the University of Belgrade (BEOU), as well as historical literature data, four bryophytes were identified as new to the flora of Serbia, namely, the mosses Fontinalis hypnoides, Leucobryum juniperoideum, Dicranum spurium and the hornwort Phaeoceros laevis. Fontinalis hypnoides is relatively recently recorded in SW Serbia, while Leucobryum juniperoideum and Dicranum spurium are known from historical collections. The hornwort Phaeoceros laevis was cited for SE Serbia in 1907, but up to now not found again.

Nevertheless, new discoveries are coming and can be expected at a regular pace (Sabovljević 2004), adding almost continually to the existing bryophyte records of Serbia (e.g.Papp et al. 2011Papp et al. , 2012aPapp et al. , b, 2013)).

MATERIALS AND METHODS
The studies on bryophyte flora of Serbia, besides field investigations, include constant activity in the revision of historical literature data and historical collections in official and unofficial collections in and outside the country.Most of the historical collections are unidentified and preserved without precise and detailed labeling.

RESULTS
Four new species are recorded for the bryophyte flora of Serbia (Fig. 1).Phaeoceros laevis (L.) Prosk. is reported as Anthoceros laevis L. in Katić (1907) for Serbia.The record was made more than 100 years ago, but is still unique.We were not able to locate the herbarium of D. Katić. Katić (1907) wrote in Serbian: on wet soil, by the river Vrla from Surdulica to Vlasina, locally quite common.

DISCUSSION
Fontinalis hypnoides is a medium-sized to robust pleurocarpous moss, often submerged.Its appearance in Serbia is not unexpected since it is present in almost all Balkan countries (Sabovljević et al. 2008).It is expected that there will be more records of this species with further bryological investigations in Serbia.
Leucobryum juniperoideum is newly reported for Serbia although all the records (five samples in total) were made in 1946 by various collectors.Two distant localities for these species infer that it could be more spread in Serbia, but overlooked.However, the genus Leucobryum, although present in Serbia, is not widespread.Thus, the records from a historical collection are also significant from the conservation point of view.In the Balkans, it is present in Slovenia, Croatia, Bulgaria and Romania (Düll et al. 1999, Sabovljević et al. 2008), so it is not strange to find it also in Serbia.As it was included into its relative L. glaucum until the 1960s, it is not surprising it is absent from all historical literature data.
Dicranum spurium is newly reported for Serbia.Its presence in Serbia is rather unexpected since it is not common in the Balkans.It is known from Slovenia, Montenegro, Bosnia-Herzegovina, Romania, and possibly Greece (Sabovljević et al. 2008).This species prefers acidic substrata and therefore needs special attention in Serbia where basic habitats are dominant.
The only hornwort reported for Serbia, Phaeoceros laevis is a forgotten species, since the only citation was made more than 100 years ago (Katić, 1907) and it has not been seen since then, although the area of its appearance was recently bryologically investigated (Papp et al. 2012c).However, it is not reported for Serbia in Pavletić (1955), Düll et al. (1999), Sabovljević (2000), Söderström et al. (2002), Sabovljević andNatcheva (2006) andROS et al. (2007).This forgotten reference is the only citation of this species in Serbia and the only record of hornworts in Serbia.There is a possibility that this species disappeared, but also that it has been overlooked due to its seasonal growth.According to Sabovljević and Natcheva (2006), it is present in the Balkans (Albania, Bulgaria, Croatia, Greece, Romania and Slovenia) and also common around the Mediterranean (Ros et al. 2007).It is taxonomically problematic and not always separated from P. carolinianus (Söderström et al. 2002), and therefore the confirmation of its presence and identity is needed.
All species mentioned, need further study of their distribution, biology and conservation.
Map of Serbia with new bryophyte records tified collection of the BEO.It was kept with dozens of other samples labeled Leucobryum.Among these samples, there were five samples of Leucobryum juniperoideum, made by three different collectors in two macro-localities: Central Serbia: -Goč Mt., Vrnjačka banja, Kraljevo; leg.M.Glišić, 08.1946(BEO s/n); Western Serbia: -Divčibare Mt., on an open slope; leg.P. Černjavski, 28.09.1946(BEO s/n); Divčibare Mt., on an open slope under Pinus sylvestris; leg.V. Lindtner, 27.09.1946(BEO s/n); Divčibare Mt., in a forest of Pinus sylvestris, leg.P. Černjavski, 28.09.1946(BEO s/n), and Divčibare Mt., in a pasture and forest of Pinus sylvestris, leg.P. Černjavski, 29.09.1946(BEO s/n) Dicranum spurium Hedw., from an unidentified collection of BEO, was recorded as new to Serbia.It was kept in a historical collection labeled as Encalypta.The sample dates from 15.09.1937, collected by P. Černjavski in central Serbia (Serbia: in a beech forest, on a southern slope ca. 2 hours walk through the gorge, south from Trstenik, schist with Leucobryum -in Serbian; su Fagetis, prope Trstenik, schistoso -in Latin) (BEO s/n).